Athyriaceae

Athyriaceae
Athyrium filix-femina
Scientific classification
Kingdom:	Plantae
Division:	Pteridophyta
Class:	Polypodiopsida /  Pteridopsida (disputed)
Order:	Polypodiales
(unranked):	Eupolypods II
Family:	Athyriaceae Alston
Genera
See text

Athyriaceae are a family of terrestrial ferns, with a cosmopolitan distribution. It is placed in the eupolypods II clade of the order Polypodiales^[1] in the class Polypodiopsida.^[2]

Genera

Athyriaceae contain the following five genera.^[1]

• Anisocampium
• Athyrium
• Cornopteris
• Deparia
• Diplazium

Athyriaceae have, in the past, included Cystopteris and Gymnocarpium, now part of Cystopteridaceae. Inversely, this family has by some been subsumed in the family Woodsiaceae, but a Woodsiaceae defined in this way may be paraphyletic if it omits the Onocleaceae and Blechnaceae (as of 2006, the evidence was not clear).^[2]

Phylogenic relationships

The following diagram for the eupolypods II, based on Lehtonen, 2011,^[3] and Rothfels & al., 2012,^[4] shows a likely phylogenic relationship between Athyriaceae and other families of the eupolypods II clade.

eupolypods II	Cystopteridaceae Rhachidosoraceae Diplaziopsidaceae Aspleniaceae Hemidictyaceae Thelypteridaceae Woodsiaceae Onocleaceae Blechnaceae Athyriaceae

This phylogenic diagram shows a possible relationship between the genera of the Athyriaceae.^[5][6]

Athyriaceae	Deparia Diplazium Anisocampium Cornopteris Athyrium

References

1. Maarten J. M. Christenhusz, Xian-Chun Zhang & Harald Schneider (2011). "A linear sequence of extant families and genera of lycophytes and ferns" (PDF). Phytotaxa. 19: 7–54.
2. Alan R. Smith; Kathleen M. Pryer; Eric Schuettpelz; Petra Korall; Harald Schneider; Paul G. Wolf (2006). "A classification for extant ferns" (PDF). Taxon. 55 (3): 705–731. doi:10.2307/25065646. Archived from the original (PDF) on 2008-02-26.
3. Samuli Lehtonen (2011). "Towards Resolving the Complete Fern Tree of Life" (PDF). PLoS ONE. 6 (10): e24851. doi:10.1371/journal.pone.0024851. PMC 3192703. PMID 22022365.
4. Carl J. Rothfels; Anders Larsson; Li-Yaung Kuo; Petra Korall; Wen- Liang Chiou; Kathleen M. Pryer (2012). "Overcoming Deep Roots, Fast Rates, and Short Internodes to Resolve the Ancient Rapid Radiation of Eupolypod II Ferns". Systematic Biology. 61 (1): 70. doi:10.1093/sysbio/sys001. PMID 22223449.
5. Eric Schuettpelz & Kathleen M. Pryer (2007). "Fern phylogeny inferred from 400 leptosporangiate species and three plastid genes" (PDF). Taxon. 56 (4): 1037–1050. doi:10.2307/25065903.
6. Yea-Chen Liu, Wen-Liang Chiou & Masahiro Kato (2011). "Molecular phylogeny and taxonomy of the fern genus Anisocampium (Athyriaceae)" (PDF). Taxon. 60 (3): 824–830.