Herichthys

Herichthys
Herichthys cyanoguttatus
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Actinopterygii
Order:	Perciformes
Family:	Cichlidae
Subfamily:	Cichlinae
Genus:	Herichthys S. F. Baird & Girard, 1854
Type species
Herichthys cyanoguttatus S. F. Baird & Girard, 1854

Herichthys is a small genus of cichlid fishes. Most are endemic to Mexico, but H. cyanoguttatus is also found in southern Texas (United States), and has been introduced to central Texas and Florida. In 2015, the genus was split, and 7 species moved into Nosferatu.

Species

There are currently 7 recognized species in this genus.^[1] Separate mitochondrial DNA analyses by Husley et al.,^[2] Rican et al.,^[3] and Oldfield et al.,^[4] support H. minckleyi as monophyletic and sister to all other Herichthys species. Moreover, molecular analysis conducted separately by López-Fernández, et al.,^[5] and De la Maza-Benignos, et al.,^[1] did not support H. tamasopoensis, H. cyanoguttatus, H. teporatus, and H. carpintis as distinct from each other. However, morphological and zoogeographical analysis by De la Maza-Benignos, et al.,^[1] confirmed the validity of these four species, as well as differential evolutionary trends. Analysis also supported the separation of Herichthys into three distinct geomorphological groups + H. minckleyi. These are, from South to North:^[1]

• Herichthys deppii (Heckel, 1840) (Nautla cichlid) ^[1][6][7]
• Herichthys tepehua De la Maza-Benignos, Ornelas-García, Lozano-Vilano, García-Ramírez & Doadrio, 2015 ^[1][7]
• Herichthys carpintis (D. S. Jordan & Snyder, 1899) (Lowland cichlid)
• Herichthys tamasopoensis Artigas Azas, 1993 (Tamasopo cichlid)
• Herichthys teporatus (Fowler, 1903) (Soto la Marina cichlid) ^[1][7]
• Herichthys cyanoguttatus (S. F. Baird & Girard, 1854) (Rio Grande cichlid)
• Herichthys minckleyi (Kornfield & J. N. Taylor, 1983) (Minkley's cichlid)

References

1. De la Maza-Benignos, M., Ornelas-García, C.P., Lozano-Vilano, M.d.L., García-Ramírez, M.E. & Doadrio, I. (2015). "Phylogeographic analysis of genus Herichthys (Perciformes: Cichlidae), with descriptions of Nosferatu new genus and H. tepehua n. sp." (PDF). Hydrobiologia. 748 (1): 201–231. doi:10.1007/s10750-014-1891-8.
2. Hulsey C.D., García , de León F.J., Sánchez-Jo Y. (2004). "Temporal diversification of Mesoamerican cichlid fishes across a major biogeographic boundary". Molecular Phylogenetics and Evolution. 31: 754–764.
3. Rıcan O.; Pialek L.; Zardoya R.; Doadrio I.; Zrzavy J. (2013). "Biogeography of the Mesoamerican Cichlidae (Teleostei: Heroini): colonization through the GAARlandia land bridge and early diversification". Journal of Biogeography. 40 (3): 579–593. doi:10.1111/jbi.12023.
4. Oldfield, R.G., Mandrekar, K., Nieves, M.X., Hendrickson, D.A., Chakrabarty, P., Swanson, B.O. & Hofmann, H.A. (2015). "Parental care in the Cuatro Ciénegas cichlid, Herichthys minckleyi (Teleostei: Cichlidae)". Hydrobiologia. 748 (1): 233–257. doi:10.1007/s10750-014-2081-4.
5. López-Fernández H.; Winemiller K.O.; Honeycutt R.L. (2010). "Multilocus phylogeny and rapid radiations in Neotropical cichlid fishes (Perciformes: Cichlidae)". Molecular Phylogenetics and Evolution. 55: 1070–1086. doi:10.1016/j.ympev.2010.02.020.
6. Hans-Joachim P., Morgenstern, R. & Schindler, I. (2014). "Cichlid fishes (Teleostei, Cichlidae) collected by Ferdinand Deppe in Mexico" (PDF). Vertebrate Zoology. 64 (1): 43–57.
7. Mcmahan C. D.; Matamoros W. A.; Piller K. R.; Chakrabarty P. (2015). "Taxonomy and systematics of the herichthyins (Cichlidae: Tribe Heroini), with the description of eight new Middle American Genera". Zootaxa. 3999 (2): 211–234. doi:10.11646/zootaxa.3999.2.3.